Node 1c: NN (Smeringopinae + Pholcinae
4 comb hairs over entire length of tarsus
(Huber & Fleckenstein 2008)
(Bruvo-Madaric et al. 2005; Astrin et al. 2007; Dimitrov et al. 2013)
All three major studies including molecules found support for a close relationship between Smeringopinae and Pholcinae.
In the plesiomorphic condition, comb hairs are restricted to the distal part of the tarsus 4: