Node 5d: NN (Subsahara Smeringopinae: Smeringopina + Smeringopus)

Two epiandrous spigots
(Huber 2012, 2013)

The male gonopore in pholcids
usually has either four epiandrous spigots (see photos at bottom of page) or none. The genera Smeringopina and Smeringopus are the only ones with two spigots.

Reduced cheliceral stridulation
(Huber 2012, 2013)

Previous analyses have interpreted cheliceral stridulation as derived within Pholcidae (e.g., Huber 2000), but this needs to be reevaluated. At least with the latest datasets, cheliceral stridulation is considered plesiomorphic for Smeringopinae, and the loss of stridulation characterizes and unites Smeringopus and Smeringopina.

Molecular data
(Dimitrov et al. 2013)

The laest molecular phylogeny of Pholcidae found good support for Smeringopus and Smeringopina. However, only two other Smeringopinae genera were included in that analysis (Holocnemus, Crossopriza).

The plesiomorphic condition is a gonopore with (usually 4, 2-6) epiandrous spigots: