Node 6: Pholcinae

pholcusMale lateral (proximal) cheliceral apophyses
(Huber 1995, 2000)

Usually weakly sclerotized projections proximally on the male chelicerae characterize a large group of spiders, originally called Pholcus-group (Huber 1995) in order to avoid Simon's Pholcinae that had a much wider scope (all pholcids except ninetines). In those taxa where the function has been studied, these apophyses interact with the retrolateral trochanter apophysis of the palp, stopping the rotation of the palp prior to insertion (Uhl et al. 1995, Huber 1995, 2001).
belisanaTarsus 4 comb hairs in single row
(Huber & Fleckenstein 2008)

Prolateral trichobothrium on tibia 1 absent
?Molecular data
12S, 28S, total evidence: Bruvo-Madaric et al. 2005; 16S+CO1, 28S: Astrin et al. 2007; Dimitrov et al. 2013)

Pholcines were consistently closely related in all studies involving genes, but in the first study the position of Spermophora was problematic, in the second study Smeringopinae were partly nested within Pholcinae.

The plesiomorphic condition is the absence of lateral (proximal) cheliceral apophyses, comb hairs in two rows, and prolateral trichobothrium present on tibia 1: